1956 | Associate of the Royal College of Science |
1956 | Bachelor of Science with Honours (BSc (Hons)) completed at the Imperial College of Science and Technology (Royal College of Science), UK |
1956 - 1957 | Research Scholar at Rothamsted Experimental Station in Harpenden, UK |
1957 - 1966 | Scientific Officer then Senior Scientific Officer at Rothamsted Experimental Station |
1961 | Doctor of Philosophy (PhD) completed at the University of London, UK |
1964 | Visiting Professor in the Faculty of Agronomy at De La Plata University in Argentina |
1966 - 1970 | Research Fellow then Senior Reseaarch Fellow at the John Curtin School of Medical Research at the Australian National University in Canberra |
1966 - 1978 | Executive Committee member of the International Committee on Taxonomy of Viruses |
1970 - 1971 | Honorary Lecturer in Virology at the University of Birmingham, UK |
1970 - 1971 | Principal Research Officer at Rothamsted Experimental Station |
1971 - | Senior Fellow at the Research School of Biological Sciences at the Australian National university |
1972 | Acting Head of the Department of Developmental Biology at the Research School of Biological Sciences |
1972 - 1975 | Honorary Associate of the Faculty of Agriculture at the University of Sydney |
1976 - 1986 | Head of the Virus Ecology Group at the Research School of Biological Sciences |
1986 - 1987 | Acting Head of the Department of Evolutionary Biology at the Research School of Biological Sciences |
1987 - 1989 | Leader of the Plant Molecular Biology Group |
1990 - | Leader of the Molecular Evolution and Systemics Group |
1993 - | Fellow of the Australian Academy of Science (FAA) |